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Grounded in the Cultural-Historical Activity Theory traditions, this qualitative case study uses the concept of relational agency to explore how residents, mentor teachers, and supervisors in a clinically-rich urban teacher residency program in the US negotiate their relationship building in supporting one another’s learning and development. Data included 17 paired-interviews (residents and mentor teachers), 6 individual supervisor interviews, and 17 individual resident interviews. Findings from content analysis suggest that relational agency enacted by participants is manifested in two types of micro-negotiations: (1) co-reflecting on the lesson or class activity; (2) bridging the theory-practice gap. The study has implications for promoting consequential collaboration across residents, mentor teachers, and supervisors in terms of developing shared truths about quality teaching and meaningful learning.