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Studies on teacher turnover have documented both negative human capital impacts (i.e., loss of experienced teachers) and financial impacts (i.e., training costs) on schools. Yet few studies attend to the organizational impacts of turnover. In this paper, we examine how teacher turnover changes instructional networks in schools—the patterns of advice seeking and support that are so critical to school improvement. We draw on constructs from social network theory, organizational theory, and theories on crisis and organizations to illuminate the different ways that turnover and external crises, such as the pandemic, influenced social networks in schools. Our longitudinal network data indicate that despite the growth in turnover rates, social relations across most levels grew closer in times of crisis.