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This research investigates the connections between need satisfaction, willingness to communicate (WTC), and quantity and quality of communication in asynchronous and synchronous learning modes. Data was collected through Experience Sampling Methodology from high school students in an online language course. Weekly surveys captured situational need satisfaction, WTC, and communication frequency. Quality of communication was assessed through writing assignments and speaking performance. Cross-lagged analyses explored temporal effects between need satisfaction, WTC, and communication behaviors. The study revealed associations between prior learning experiences, perceptions, and subsequent engagements over time. It provides valuable insights into contextual factors influencing students' communication behaviors within different learning modes and content units.