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We use the concept of agency to explore university-district partnerships formed to advance equity-centered school leader preparation. Using Cultural-Historical Activity Theory (CHAT), we analyze how one urban school district and its two university partners navigated institutional differences during a multi-year, grant-funded initiative focused on leadership for equity. Data analysis addresses three forms of agency: individual agency that included boundary-challenging actions, relational agency that helped partners balance institutional autonomy with collaborative expansion, and collective agency that gave rise to shared practices. We conceptualize the partnership as a third space, and demonstrate how productive tension resolution and collaborative meaning-making expanded the partners’ capacities to advance equity beyond what any institution could have achieved independently.