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Nonprofit participation in the form of giving and volunteering has long been viewed as the key building blocks of participatory democracy. Yet prior research has rarely treated nonprofit participation as a distinctive form of civic participation. The current study extends the Communication Mediation Model (CMM) to examine nonprofit-specific participation behaviors within local communication possesses. It uses structural equation modeling to investigate the paths of influence among community attachment variables, nonprofit-specific media use, nonprofit-specific discussion, and individuals’ volunteering and giving behaviors. Results highlight the importance of nonprofit-specific discussion in (1) directly promoting both giving and volunteering behaviors, (2) mediating the influence of nonprofit-specific traditional media use, and (3) translating community attachment into greater nonprofit involvement. Findings also identify a different set of community and communication mechanisms significantly associated with nonprofit giving versus volunteering. Practical implications are discussed in the context of leveraging communication processes to promote local nonprofit participation.