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Using novel methodological approaches to assess how the heterogeneous landscapes created by complex agroecosystems can host a relevant share of functional biodiversity, the communication will present the results obtained in several recent case studies in Catalonia, Spain (1956-2009) and Costa Rica (1986-2014) that statistically correlate landscape ecology indicators accounted in a set of cells of a grid in past and present land cover digital maps with the biodiversity observations currently made in the same cells. Despite founding significant differences between Mediterranean and Tropical landscapes, the results confirm that biodiversity is located both in nature protected areas and the heterogeneous landscape mosaics of complex agroecosystems. Conversely, industrial monocultures as well as rural abandonment have a negative ecological impact, whereas wildlife-friendly agroecology farming plays a vital role either to keep farm-associated biodiversity or to provide ecological connectivity among nature protected areas.