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Mathematics content courses centering on student thinking can play a significant role in fostering positive affect in prospective teachers (PTs) (Auslander, Hart, Carothers, et al., 2019). This study explores two prospective elementary teachers’ beliefs about mathematics teaching and learning and their performance on a mixed fraction task that involves analyzing and sequencing student work. Despite the level of their conceptual and procedural understanding of the concept, PTs’ performance in the task-based interview contrasted with the self-reported views. These findings highlighted the need for mathematics teacher education programs to address the complexity of the interplay among mathematics beliefs, prior learning experience, and mathematics proficiency to support PTs in developing productive beliefs that they can enact to foster meaningful mathematics learning for their future students.