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This convergent mixed methods study examined pre-service teachers’ beliefs about in-service teacher collaboration, and how those beliefs influenced their collaboration in four-member groups during design and delivery of an instructional project. Participants (N=24) were in their first semester of under-graduate teacher preparation. Both quantitative and qualitative data were collected. Integrated analysis indicated all participants believed in-service collaboration involves cooperative activities of joint planning, and sharing ideas and resources (simple beliefs). Very few participants also believed in-service collaboration includes activities of collective assessment and evaluation (expanded beliefs). Findings suggest pre-service beliefs about in-service collaboration impacts students’ collaborative efforts and outcomes. Implications for practice and research indicates understanding factors influencing pre-service collaboration may help teacher preparation better address needs of students.